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Acknowledgements This work was supported by National Key Research and Development Program of China (2022YFA1105300), National Natural Science Foundation of China (82273241, 82102921, 82203822), China Postdoctoral Science Foundation (2023M744049), Guangdong Basic and Applied Basic Research Foundation (2023B1515040030, 2023A1515010547, 2021A1515111001), Sanming Project of Medicine in Shenzhen (SZSM202211017), Chih Kuang Scholarship for Outstanding Young Physician-Scientists of Sun Yat-sen University Cancer Center (CKS-SYSUCC-2023001)

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